iTOL version:	7.5
Tree of Life:	version 1.0 (Ciccarelli FD, et al., Science.; 2006 Mar 3. 311(5765): 1283-7)
Publications:	Letunic I and Bork P (2006) Bioinformatics 23(1):127-8 Interactive Tree Of Life (iTOL): an online tool for phylogenetic tree display and annotation (abstract, full article) Letunic I and Bork P (2011) Nucleic Acids Res doi: 10.1093/nar/gkr201 Interactive Tree Of Life v2: online annotation and display of phylogenetic trees made easy (abstract, full article) Letunic I and Bork P (2016) Nucleic Acids Res doi: 10.1093/nar/gkw290 Interactive Tree Of Life (iTOL) v3: an online tool for the display and annotation of phylogenetic and other trees (abstract, full article) Letunic I and Bork P (2019) Nucleic Acids Res doi: 10.1093/nar/gkz239 Interactive Tree Of Life (iTOL) v4: recent updates and new developments (full article) Letunic I and Bork P (2021) Nucleic Acids Res doi: 10.1093/nar/gkab301 Interactive Tree Of Life (iTOL) v5: an online tool for phylogenetic tree display and annotation (full article) Letunic I and Bork P (2024) Nucleic Acids Res doi: 10.1093/nar/gkae268 Interactive Tree of Life (iTOL) v6: recent updates to the phylogenetic tree display and annotation tool (full article)

It is with immense sadness that I share the news of the passing on 16 January 2026, of my colleague and friend Peer Bork.

Peer was a visionary, and his career was far too rich and intense to be summarized in just a few lines. His work on phylogenetics and the reconstruction of the Tree Of Life from fully sequenced genomes led to the initial idea and early development of iTOL. Without his leadership and support, it would never have grown into what it is.

You will be remembered forever.

Local installations and customization of iTOL

If you are working with sensitive data, or would be interested in running a customized version of iTOL in your own organization, please contact us. We can provide full integration with your exiting infrastructure, or complete customization of iTOL to fit your specific needs.

Recent changes and new features

The complete list of iTOL updates is available at: the version history page

Feedback and contact

We hope you find iTOL user-friendly and easy to use. However, we encourage anyone who:

• has problems using or understanding iTOL's functions
• cannot upload a custom tree or a dataset/annotation file
• encounters a server error

to contact us. You can also use the following form to send a quick question or feedback: